Lions live in groups of related females called prides, which may comprise several to as many as 40 individuals, including adults, sub-adults (two to four years old) and cubs, plus one or more resident males. Abundance of prey availability plays a significant role in the size of a lion pride. Pride mates associate in sub-groups within the pride. Females stay in their mothers’ prides for life, unless food scarcity forces them out. Young males are driven from their prides when they grow large enough to compete with the dominant males. Young males join in coalitions, usually with brothers and cousins, and search for a pride to take over. Males entering a new pride will kill all cubs that cannot run from them. Adult males that are fortunate enough achieve residency within a pride hold tenure for an average of two years, often leaving due to eviction by another coalition of males. In India, female and male lions live apart, joining only to mate.